Publications

(Also see publications on Google Scholar, Research Gate, and CORE.)

Northern development and Rangifer risks: A review of the impacts of resource extraction for caribou and reindeer Lessard É, Walker PD, Vander Wal E._{10.1139/er-2025-0182}

Landscapes of fearlessness revealed through hormonal responses to putatively risky places Newediuk L, Jesmer BR, Mastromonaco G, Vander Wal E._{10.1093/beheco/arag004}

Addled by antlers: Synchronous disruption to female caribou antler phenology Love AE, Fox JA, Hendrix JG, Jackson S, Ferraro K, Vander Wal E, Webber QMR._{10.1002/ecs2.70484}

Movement-integrated habitat selection reveals wolves balance ease of travel with human avoidance in a risk–reward trade-off Kingdon KA, Prokopenko CM, Dupont DLJ, Turner J, Robitaille AL, Wiens J, Harriman VB, Vander Wal E._{10.1111/1365-2664.70031}

One size does not fit all: A novel approach for determining the Realised Viewshed Size for remote camera traps Carswell BM, Avgar T, Street GM, Boyle SP, Vander Wal E._{10.1111/2041-210X.70008}

Consistent Individual Differences and Plasticity in Migration Behaviour of Three North American Ungulates Laforge MP, Vander Wal E, Webber QMR, Geremia C, Kauffman MJ, McWhirter DE, Middleton A, Mong TW, Monteith KL, Ortega AC, Sawyer H, Merkle JA._{10.1111/ele.70101}

Sociality and movement change through space and time: Implications for anti-predator strategies in caribou Atkinson M, Love AE, Hendrix JG, Kusch JM, Ellington EH, Vander Wal E, Webber QMR._{10.1139/cjz-2024-0092}

Intrinsic and environmental drivers of pairwise cohesion in wild Canis social groups Benson JF, Keiter DA, Mahoney PJ, Allen BL, Allen L, Álvares F, Anderson ML, Barber-Meyer SM, Barocas A, Beasley JC, Behrendorff L, Belant JL, Beyer DE, Boitani L, Borg BL, Boutin S, Boydston EE, Brown JL, Bump JK, Cepek JD, Chamberlain MJ, Chenaux-Ibrahim YM, Cherry SG, Ćirović D, Ciucci P, Cluff HD, Cooper SM, Crooks KR, Dupont DLJ, Fisher RN, Fortin D, Gable TD, García E, Geffen E, Gehrt SD, Gillingham M, Heard DC, Hebblewhite M, Hinton JW, Homkes AT, Howden CG, Huber D, Jackson PJ, Joly K, Kelly A, Kelly MJ, Kingdon KA, Kulkarni A, Kusak J, Kuzyk GW, Lake BC, Llaneza L, López-Bao JV, MacNulty DR, McLaren AAD, McLoughlin PD, Merrill EH, Mills KJ, Mitchell N, Moore SA, Mumma MA, Murray MH, Musiani M, Nakamura M, Neilson EW, Neufeld LM, Newsome TM, Oakleaf JK, Palacios V, Perdicas MM, Perry T, Petroelje TR, Piper CB, Prokopenko CM, Prugh LR, Riley SPD, Rio-Maior H, Roffler GH, Rollins D, Sand H, Schmiegelow FKA, Seip DR, Sorum MS, St. Clair CC, Steenweg R, Strohbach MW, Tatler J, Thaker M, Thompson CA, Turner JW, Vanak AT, Vander Wal E, Wabakken P, Walter SE, Webster SC, Wheeldon TJ, Wikenros C, Windels SK, Young JK, Zabihi-Seissan S, Zimmermann B, Patterson BR._{10.1002/ecy.4492}

Bridging reproductive management and ecosystem function: Behavioural pathways linking contraception to zoogeochemistry Funk KR, Ferraro KM, Schulte-Hostedde AI, Wal EV._{10.1139/cjz-2025-0076}

Social and seasonal risk assessment by woodland caribou: Responses to habitat openness, fire, and roads Hendrix JG, Brown CD, Gosse J, Vander Wal E._{10.1139/cjz-2024-0150}

Density-dependent network structuring within and across wild animal systems Albery GF, Becker DJ, Firth JA, De Moor D, Ravindran S, Silk M, Sweeny AR, Vander Wal E, Webber Q, Allen B._{10.1038/s41559-025-02843-z}

Zoogeochemical niche construction: How animal-mediated biogeochemistry affects evolution Ferraro KM, Leroux SJ, Bradford MA, Schmitz OJ, Vander Wal E._{10.1016/j.tree.2025.07.009}

Body condition of translocated African elephants (Loxodonta africana) in a semi-arid environment: Effects of seasonality, age-sex, and management interventions Funk KR, Brown LR, Vander Wal E._{10.1016/j.jnc.2025.126959}

Quantifying elemental diversity to study landscape ecosystem function Mcleod AM, Leroux S, Little CL, Massol F, Vander Wal E, Wiersma YF, Gounand I, Loeuille N, Harvey E._{10.1016/j.tree.2024.09.007}

Expanding theory, methodology and empirical systems at the spatial–social interface Albery GF, Webber QMR, Farine D, Picardi S, Vander Wal E, Manlove KR._{10.1098/rstb.2022.0534}

Faithful pals and familiar locales: Differentiating social and spatial site fidelity during reproduction Hendrix JG, Robitaille AL, Kusch JM, Webber QMR, Vander Wal E._{10.1098/rstb.2022.0525}

Friends because of foes: Synchronous movement within predator–prey domains Prokopenko CM, Ellington EH, Robitaille A, Aubin JA, Balluffi-Fry J, Laforge M, Webber QMR, Zabihi-Seissan S, Vander Wal E._{10.1098/rstb.2023.0374}

Effects of the social environment on movement-integrated habitat selection Webber Q, Prokopenko C, Kingdon K, Turner J, Vander Wal E._{10.1186/s40462-024-00502-9}

The biogeochemical boomerang: Site fidelity creates nutritional hotspots that may promote recurrent calving site reuse Ferraro KM, Albrecht D, Hendrix JG, Wal EV, Schmitz OJ, Webber QMR, Bradford MA._{10.1111/ele.14491}

Variation in spatiotemporal activity may reduce competitive interactions between invasive wild pigs ( Sus Scrofa ) and native mammal species Carswell BM, Boyle SP, Brook RK, Van Beest FM, Vander Wal E._{10.1139/cjz-2022-0145}

Associations between glucocorticoids and habitat selection reflect daily and seasonal energy requirements Newediuk L, Mastromonaco GF, Vander Wal E._{10.1186/s40462-024-00475-9}

Environmental, social and morphological drivers of fission–fusion dynamics in a social ungulate Le Goff M, Hendrix JG, Webber QM, Robitaille AL, Vander Wal E.

Animal-vectored nutrient flows across resource gradients influence the nature of local and meta-ecosystem functioning Rizzuto M, Leroux SJ, Schmitz OJ, Vander Wal E, Wiersma YF, Heckford TR._{10.1016/j.ecolmodel.2023.110570}

Post-rut aggression in caribou: Limited evidence for interference competition but strong evidence for male harassment Webber QM, Vander Wal E._{10.1163/1568539X-bja10276}

The adaptive value of density-dependent habitat specialization and social network centrality Webber QM, Laforge MP, Bonar M, Vander Wal E._{10.1038/s41467-024-48657-8}

Functional Response to Cumulative Effects as an Effective Tool for Wildlife Management Kingdon KA, Stewart FEC, Wiersma YF, Vander Wal E._{10.1007/s40823-023-00094-x}

Death comes for us all: Relating movement-integrated habitat selection and social behavior to human-associated and disease-related mortality among gray wolves Turner JW, Prokopenko CM, Kingdon KA, Dupont DLJ, Zabihi-Seissan S, Vander Wal E._{10.1007/s00442-023-05426-6}

Behavioural ecology at the spatial-social interface Webber QMR, Albery GFF, Farine DRR, Pinter-Wollman N, Sharma N, Spiegel O, Vander Wal E, Manlove K._{10.1111/brv.12934}

The problem and promise of scale in multilayer animal social networks Robitaille AL, Webber QMR, Turner JW, Vander Wal E._{10.1093/cz/zoab068}

Coat color mismatch improves survival of a keystone boreal herbivore: Energetic advantages exceed lost camouflage Kennah JL, Peers MJL, Vander Wal E, Majchrzak YN, Menzies AK, Studd EK, Boonstra R, Humphries MM, Jung TS, Kenney AJ, Krebs CJ, Boutin S._{10.1002/ecy.3882}

Protection, energetic assistance, or social perks: How do beluga offspring benefit from allocare? Aubin JA, Michaud R, Vander Wal E._{10.1111/mms.12957}

Uncertainty during migration: The role of predictable resources in unpredictable conditions in determining movement in Woodland Caribou (Rangifer tarandus) Dedeban E, Webber QM, Laforge MP, Robitaille AL, Vander Wal E._{10.1093/jmammal/gyad081}

Plasticity and repeatability in spring migration and parturition dates with implications for annual reproductive success Laforge MP, Webber QMR, Vander Wal E._{10.1111/1365-2656.13911}

Wolf spatial behavior promotes encounters and kills of abundant prey Zabihi-Seissan S, Prokopenko CM, Vander Wal E._{10.1007/s00442-022-05218-4}

Ecoregion and community structure influences on the foliar elemental niche of balsam fir (Abies balsamea (L.) mill.) and white birch (Betula papyrifera marshall) Heckford TR, Leroux SJ, Vander Wal E, Rizzuto M, Balluffi-Fry J, Richmond IC, Wiersma YF._{10.1002/ece3.9244}

Predicting the individual identity of non-invasive faecal and hair samples using biotelemetry clusters Newediuk L, Vander Wal E._{10.1007/s42991-021-00173-8}

In defense of elemental currencies: Can ecological stoichiometry stand as a framework for terrestrial herbivore nutritional ecology? Balluffi-Fry J, Leroux SJ, Champagne E, Vander Wal E._{10.1007/s00442-022-05176-x}

Repeatability is the first step in a broader hypothesis test: A comment on Stuber et al Vander Wal E, Webber QMR, Laforge MP._{10.1093/beheco/arac005}

What do caribou eat? A review of the literature on caribou diet Webber QMR, Ferraro KM, Hendrix JG, Vander Wal E._{10.1139/cjz-2021-0162}

The relevance of genetic structure in ecotype designation and conservation management Stronen A V, Norman AJ, Vander Wal E, Paquet PC._{10.1111/eva.13339}

Spatially explicit correlates of plant functional traits inform landscape patterns of resource quality Heckford TR, Leroux SJ, Vander Wal E, Rizzuto M, Balluffi-Fry J, Richmond IC, Wiersma YF._{10.1007/s10980-021-01334-3}

Individual differences in habitat selection mediate landscape level predictions of a functional response Newediuk L, Prokopenko CM, Vander Wal E._{10.1007/s00442-021-05098-0}

Conceptual and methodological advances in habitat-selection modeling: Guidelines for ecology and evolution Northrup JM, Vander Wal E, Bon M, Fieberg J, Laforge MP, Leclerc M, Prokopenko CM, Gerber BD._{10.1002/eap.2470}

Individual snowshoe hares manage risk differently: Integrating stoichiometric distribution models and foraging ecology Richmond IC, Balluffi-Fry J, Vander Wal E, Leroux SJ, Rizzuto M, Heckford TR, Kennah JL, Riefesel GR, Wiersma YF, Pauli J._{10.1093/jmammal/gyab130}

Optimal prey switching: Predator foraging costs provide a mechanism for functional responses in multi-prey systems Prokopenko CM, Avgar T, Ford A, Vander Wal E._{10.1002/ecy.3928}

Solving the sample size problem for resource selection functions Street GM, Potts JR, Borger L, Beasley JC, Demarais S, Fryxell JM, McLoughlin PD, Monteith KL, Prokopenko CM, Ribeiro MC, Rodgers AR, Strickland BK, van Beest FM, Bernasconi DA, Beumer LT, Dharmarajan G, Dwinnell SP, Keiter DA, Keuroghlian A, Newediuk LJ, Oshima JEF, Rhodes O Jr., Schlichting PE, Schmidt NM, Vander Wal E._{10.1111/2041-210X.13701}

On the marginal value of swimming in woodland caribou Webber QMR, Hendrix JG, Robitaille AL, Vander Wal E._{10.1002/ecy.3491}

Forage stoichiometry predicts the home range size of a small terrestrial herbivore Rizzuto M, Leroux SJ, Vander Wal E, Richmond IC, Heckford TR, Balluffi-Fry J, Wiersma YF._{10.1007/s00442-021-04965-0}

Context-dependent group size: Effects of population density, habitat, and season Webber QMR, Vander Wal E._{10.1093/beheco/arab070}

Temporal variation and its drivers in the elemental traits of four boreal plant species Richmond IC, Leroux SJ, Heckford TR, Vander Wal E, Rizzuto M, Balluffi-Fry J, Kennah JL, Wiersma YF._{10.1093/jpe/rtaa103}

Coyote (Canis latrans) diet and spatial co-occurrence with woodland caribou (Rangifer tarandus caribou) Huang RKK, Webber QMR, Laforge MP, Robitaille AL, Bonar M, Balluffi-Fry J, Zabihi-Seissan S, Vander Wal E._{10.1139/cjz-2020-0253}

Sociopolitical factors drive conservation planning timelines: A canadian case study with global implications Newediuk L, Ethier JP, Boyle SP, Aubin JA, Balluffi-Fry J, Dedeban E, Laforge MP, Prokopenko CM, Turner JW, Webber QMR, Vander Wal E._{10.1016/j.biocon.2021.109091}

Tracking snowmelt to jump the green wave: Phenological drivers of migration in a northern ungulate Laforge MP, Bonar M, Vander Wal E._{10.1002/ecy.3268}

Social environment: Trait, context and agent for selection in a meta-population Vander Wal E._{10.1111/1365-2656.13400}

Prospective evolutionary drivers of allocare in wild belugas Aubin JA, Michaud R, Wal EV._{10.1163/1568539X-bja10094}

Integrating plant stoichiometry and feeding experiments: State-dependent forage choice and its implications on body mass Balluffi-Fry J, Leroux SJ, Wiersma YF, Richmond IC, Heckford TR, Rizzuto M, Kennah JL, Vander Wal E._{10.1007/s00442-021-05069-5}

Quantity-quality trade-offs revealed using a multiscale test of herbivore resource selection on elemental landscapes Balluffi-Fry J, Leroux SJ, Wiersma YF, Heckford TR, Rizzuto M, Richmond IC, Vander Wal E._{10.1002/ece3.6975}

Herbivore impacts on carbon cycling in boreal forests Leroux SJ, Wiersma YF, Vander Wal E._{10.1016/j.tree.2020.07.009}

Is less more? A commentary on the practice of “metric hacking” in animal social network analysis Webber QMR, Schneider DC, Vander Wal E._{10.1016/j.anbehav.2020.08.011}

Geometry of the ideal free distribution: Individual behavioural variation and annual reproductive success in aggregations of a social ungulate Bonar M, Lewis KP, Webber QMR, Dobbin M, Laforge MP, Vander Wal E._{10.1111/ele.13563}

Divergent estimates of herd-wide caribou calf survival: Ecological factors and methodological biases Ellington EH, Lewis KP, Koen EL, Vander Wal E._{10.1002/ece3.6553}

Individual responses to novel predation risk and the emergence of a landscape of fear Perry TA, Laforge MP, Vander Wal E, Knight TW, McLoughlin PD._{10.1002/ecs2.3216}

The ecology of individual differences empirically applied to space-use and movement tactics Webber QMR, Laforge MP, Bonar M, Robitaille AL, Hart C, Zabihi-Seissan S, Vander Wal E._{10.1086/708721}

Heterogeneity in social network connections is density-dependent: Implications for disease dynamics in a gregarious ungulate Webber QMR, Vander Wal E._{10.1007/s00265-020-02860-x}

Biologging special feature Borger L, Bijleveld AI, Fayet AL, Machovsky-Capuska GE, Patrick SC, Street GM, Vander Wal E._{10.1111/1365-2656.13163}

Density dependence and eco-evolutionary dynamics of animal social networks: A comment on shizuka and johnson Vander Wal E, Webber QMR._{10.1093/beheco/arz150}

Patterns and potential drivers of intraspecific variability in the body C, N, and P composition of a terrestrial consumer, the snowshoe hare (Lepus americanus) Rizzuto M, Leroux SJ, Vander Wal E, Wiersma YF, Heckford TR, Balluffi-Fry J._{10.1002/ece3.5880}

Conducting social network analysis with animal telemetry data: Applications and methods using spatsoc Robitaille AL, Webber QMR, Vander Wal E._{10.1111/2041-210X.13215}

Understanding habitat co-occurrence and the potential for competition between native mammals and invasive wild pigs (Sus scrofa) at the northern edge of their range O’Brien P, Wal EV, Koen EL, Brown CD, Guy J, van Beest FM, Brook RK._{10.1139/cjz-2018-0156}

Space use and social association in a gregarious ungulate: Testing the conspecific attraction and resource dispersion hypotheses Peignier M, Webber QMR, Koen EL, Laforge MP, Robitaille AL, Vander Wal E._{10.1002/ece3.5071}

Trends and perspectives on the use of animal social network analysis in behavioural ecology: A bibliometric approach Webber QMR, Vander Wal E._{10.1016/j.anbehav.2019.01.010}

Reproductive ecology of recently established wild pigs in canada Koen EL, Vander Wal E, Kost R, Brook RK._{10.1674/0003-0031-179.2.275}

Wildlife biology, big data, and reproducible research Lewis KP, Vander Wal E, Fifield DA._{10.1002/wsb.847}

Consistent individual differences and population plasticity in network-derived sociality: An experimental manipulation of density in a gregarious ungulate O’Brien PP, Webber QMR, Vander Wal E._{10.1371/journal.pone.0193425}

Implementing a novel movement-based approach to inferring parturition and neonate caribou calf survival Bonar M, Ellington EH, Lewis KP, Vander Wal E._{10.1371/journal.pone.0192204}

An evolutionary framework outlining the integration of individual social and spatial ecology Webber QMR, Vander Wal E._{10.1111/1365-2656.12773}

Stoichiometric distribution models: Ecological stoichiometry at the landscape extent Leroux SJ, Vander Wal E, Wiersma YF, Charron L, Ebel JD, Ellis NM, Hart C, Kissler E, Saunders PW, Moudra L, Tanner AL, Yalcin S._{10.1111/ele.12859}

Does prey density predict characteristics of primiparity in a solitary and specialized predator, the Canada lynx (Lynx canadensis)? Reynolds JJ, Vander Wal E, Adams BK, Curran RM, Doucet CM._{10.1139/cjz-2016-0269}

Observation of a p [less than] 10\(^-\)9 life-history event: Implications of record-late caribou birth on ungulate reproductive ecology and field studies Bonar M, Laforge MP, Vander Wal E._{10.1139/cjz-2016-0245}

Quantifying consistent individual differences in habitat selection Leclerc M, Wal EV, Zedrosser A, Swenson JE, Kindberg J, Pelletier F._{10.1007/s00442-015-3500-6}

Dyadic associations and individual sociality in bighorn ewes Vander Wal E, Gagne-Delorme A, Festa-Bianchet M, Pelletier F._{10.1093/beheco/arv193}

Process-focussed, multi-grain resource selection functions Laforge MP, Vander Wal E, Brook RK, Bayne EM, McLoughlin PD._{10.1016/j.ecolmodel.2015.03.003}

Born to be wild? Response of an urban exploiter to human-modified environment and fluctuating weather conditions Turgeon G, Vander Wal E, Masse A, Pelletier F._{10.1139/cjz-2014-0263}

Diurnal and nocturnal activity patterns of invasive wild boar (Sus scrofa) in Saskatchewan, Canada Stolle K, van Beest FM, Vander Wal E, Brook RK._{10.22621/cfn.v129i1.1670}

Density-dependent habitat selection and partitioning between two sympatric ungulates van Beest FM, McLoughlin PD, Vander Wal E, Brook RK._{10.1007/s00442-014-2978-7}

Evolutionary perspectives on wildlife disease: Concepts and applications Vander Wal E, Garant D, Pelletier F._{10.1111/eva.12179}

Applying evolutionary concepts to wildlife disease ecology and management Vander Wal E, Garant D, Calme S, Chapman CA, Festa-Bianchet M, Millien V, Rioux-Paquette S, Pelletier F._{10.1111/eva.12168}

Density dependence in social behaviour: Home range overlap and density interacts to affect conspecific encounter rates in a gregarious ungulate Vander Wal E, Laforge MP, McLoughlin PD._{10.1007/s00265-013-1652-0}

Increasing density leads to generalization in both coarse-grained habitat selection and fine-grained resource selection in a large mammal van Beest FM, Uzal A, Vander Wal E, Laforge MP, Contasti AL, Colville D, McLoughlin PD._{10.1111/1365-2656.12115}

Sex-based differences in the adaptive value of social behavior contrasted against morphology and environment Vander Wal E, Festa-Bianchet M, Réale D, Coltman DW, Pelletier F._{10.1890/14-1320.1}

Juxtaposition between host population structures: Implications for disease transmission in a sympatric cervid community Vander Wal E, Edye I, Paquet PC, Coltman DW, Bayne E, Brook RK, Andres JA._{10.1111/eva.12065}

Targeting hunter distribution based on host resource selection and kill sites to manage disease risk Dugal CJ, van Beest FM, Vander Wal E, Brook RK._{10.1002/ece3.788}

Factors driving variation in movement rate and seasonality of sympatric ungulates van Beest FM, Wal EV, Stronen AV, Brook RK._{10.1644/12-MAMM-A-080.1}

Evaluating use of cattle winter feeding areas by elk and white-tailed deer: Implications for managing bovine tuberculosis transmission risk from the ground up Brook RK, Vander Wal E, van Beest FM, McLachlan SM._{10.1016/j.prevetmed.2012.07.017}

Density-dependent effects on group size are sex-specific in a gregarious ungulate Vander Wal E, van Beest FM, Brook RK._{10.1371/journal.pone.0053777}

Identifying hidden sinks in growing populations from individual fates and movements: The feral horses of sable island Contasti AL, Van Beest FM, Vander Wal E, McLoughlin PD._{10.1002/jwmg.625}

Context-dependent social behaviour: Testing the interplay between season and kinship with raccoons Robert K, Garant D, Vander Wal E, Pelletier F._{10.1111/jzo.12027}

Temporal variation in site fidelity: Scale-dependent effects of forage abundance and predation risk in a non-migratory large herbivore van Beest FM, Vander Wal E, Stronen AV, Paquet PC, Brook RK._{10.1007/s00442-013-2647-2}

Evolutionary rescue in vertebrates: Evidence, applications and uncertainty Vander Wal E, Garant D, Festa-Bianchet M, Pelletier F._{10.1098/rstb.2012.0090}

Effects of phenology and sex on social proximity in a gregarious ungulate Vander Wal E, Paquet PC, Messier F, McLoughlin PD._{10.1139/cjz-2012-0237}

Influence of landscape and social interactions on transmission of disease in a social cervid Vander Wal E, Paquet PC, Andres JA._{10.1111/j.1365-294X.2011.05431.x}

An individual-based quantitative approach for delineating core areas of animal space use Vander Wal E, Rodgers AR._{10.1016/j.ecolmodel.2011.10.006}

Sex-based differences in density-dependent sociality: An experiment with a gregarious ungulate Vander Wal E, Yip H, McLoughlin PD._{10.1890/11-0020.1}

Spatial and temporal factors influencing sightability of elk Vander Wal E, McLoughlin PD, Brook RK._{10.1002/jwmg.208}

Seasonal shifts in habitat selection of a large herbivore and the influence of human activity McLoughlin PD, Vander Wal E, Lowe SJ, Patterson BR, Murray DL._{10.1016/j.baae.2011.09.001}

Considering ecological dynamics in resource selection functions McLoughlin PD, Morris DW, Fortin D, Vander Wal E, Contasti AL._{10.1111/j.1365-2656.2009.01613.x}

Designating seasonality using rate of movement Wal EV, Rodgers AR._{10.2193/2008-137}